http://steamcommunity.com/games/593110/ ... 4614253751
The main difference is that the Steam community is no longer going to be involved in the decision making if a title is worthy to be released on Steam or not and instead of the one time publishing fee, developers are going to be required to pay for each title and the fee can be as low as 100 USD and as high as 5000 USD per title. At this point Valve didn't made a decision on this, they are still talking to developers to get feedback and set the final price.
Of course what it really means we can't know for sure until Steam Direct actually replace Steam Greenlight and Valve set the final price for publishing fee but what it looks like, it means better quality control, better, more polished titles released in the store which is generally a good thing for gamers, and for indie developers it means their titles need to be more polished and better designed (than usual) to make it in to the store and also need to be dedicated to provide great and fun content worthy to pay the potentially higher per title fee.
The bottom line is, a supporting community is no longer going to be enough and "to pay for support (offer steam key for vote)" is no longer going to be an option to get a game published on Steam.